Publications

2025

Reed, S. P., Royo, A. A., Carson, W. P., Olmsted, C. F., Frelich, L. E., & Reich, P. B. (2025). Multiple disturbances, multiple legacies: Fire, canopy gaps and deer jointly change the forest seed bank. Journal of Ecology, 113, 353–370. PDF

Tremblay, J. P., Champagne, E., Gill, R., Iijima, H., & Royo, A. A. (2025). Deer impacts on forest conservation. In M. Melletti & S. Focardi (Eds.), Deer of the World – Ecology, Conservation, and Management (pp. 913–924). Springer Press. https://doi.org/10.1007/978-3-031-17756-9

Vickers, L. A., Royo, A. A., Stout, S. L., Ristau, T., Oldland, W. K., Knapp, B. O., Knapp, L. S. P., & Hille, A. (2025). Evaluating the extent and severity of black cherry regeneration declines in eastern forests and exploring potential causes. In B. L. Conkling & K. Pandit (Eds.), Forest Health Monitoring: Evaluation Monitoring Project Summaries 2023 (Gen. Tech. Rep. WO-105a, pp. 9–16). U.S. Department of Agriculture, Forest Service. PDF

2024

Adams, B. T., Royo, A. A., Kern, C. C., Bronson, D. R., Matthews, S. N., Gougherty, A. V., Prasad, A. M., Iverson, L. R., Rehm, E. M., Raymond, P., Périé, C., Miesel, J. R., & McGraw, A. M. (2024). Identifying climatically-compatible seedlots for the eastern US: Building the predictive tools and knowledge to enable forest assisted migration. Frontiers in Forests and Global Change, 7. PDF

Bower, A. D., Frerker, K. L., Pike, C. C., Labonte, N. R., Palik, B. J., Royo, A. A., Anderson, S. M., Ferreira, A. R., & Brandt, L. A. (2024). A practical framework for applied forestry assisted migration. Frontiers in Forests and Global Change, 7. PDF

Glover, L. M., Volponi, S. N., Royo, A. A., & Johnstone-Yellin, T. L. (2024). Effect of herbivory on the nutritional value of six hardwood species for white-tailed deer in the Allegheny Forest. Journal of Fish and Wildlife Management, 15, 183–194. PDF

Hutchinson, T. F., Adams, B. T., Dickinson, M. B., Heckel, M., Royo, A. A., & Thomas-Van Gundy, M. A. (2024). Sustaining eastern oak forests: Synergistic effects of fire and topography on vegetation and fuels. Ecological Applications, 34, e2948. PDF

Locke, E., Marsh, M., O’Brien, M., Rowan, E., Rivera, M., Pinchot, C., Ristau, T. E., & Royo, A. A. (2024). Deer browsing effects on herbaceous layer diversity following shelterwood harvests. In M. Thomas-Van Gundy & M. B. Adams (Comps.), Proceedings of the 22nd Central Hardwood Forest Conference: Managing future forests today (Gen. Tech. Rep. NRS-219, pp. 133–148). U.S. Department of Agriculture, Forest Service, Northern Research Station. PDF

Rowan, E., Slaughter, M., Dzemyan, J., Nelson, B., & Royo, A. A. (2024). The Kinzua Quality Deer Cooperative: Balancing white-tailed deer populations and their browsing impact to forests over two decades of local land stewardship. In M. Thomas-Van Gundy & M. B. Adams (Eds.), Proceedings of the 22nd Central Hardwood Forest Conference: Managing future forests today (Gen. Tech. Rep. NRS-219, pp. 13–23). U.S. Department of Agriculture, Forest Service, Northern Research Station. PDF

2023

Brault, B., Tremblay, J. P., Thiffault, N., Royo, A. A., & Côté, S. D. (2023). Successful forest restoration using plantations: How neighboring vegetation drives browsing pressure and tree growth. Forest Ecology and Management, 549, 121458. PDF

Champagne, E., Royo, A. A., Tremblay, J. P., & Raymond, P. (2023). Tree assisted migration in a browsed landscape: Developing an assessment of susceptibility. In C. C. Kern & Y. L. Dickinson (Eds.), Proceedings of the first biennial Northern Hardwood Conference 2021: Bridging science and management for the future (Gen. Tech. Rep. NRS-P-211, pp. 275–279). U.S. Department of Agriculture, Forest Service, Northern Research Station. PDF

Knight, K. S., Flower, C. E., Kilgore, J. S., Royo, A. A., Fox, T. D., Kappler, R. H., Hoven, B. M., Wigal, J. S., Bernard, J. D., & Long, R. P. (2023). Ash tree decline and mortality in Ohio and the Allegheny National Forest in Pennsylvania. In K. M. Potter & B. L. Conkling (Eds.), Forest Health Monitoring: National status, trends, and analysis 2022 (Gen. Tech. Rep. SRS-273, pp. 139–147). U.S. Department of Agriculture, Forest Service, Southern Research Station. PDF

Pinchot, C., Royo, A. A., Stanovick, J., Schlarbaum, S., Sharp, A., & Anagnostakis, S. (2023). Browsing by deer reduces growth and competitive ability of planted backcross chestnut seedlings despite also favoring slower-growing competing species. Forest Ecology and Management, 523, 120481. PDF

Refsland, T. K., Adams, B., Bronson, D., Kern, C. C., Marquardt, P., McGraw, A. M., Royo, A. A., & Miesel, J. R. (2023). Trait-mediated effects of plant-soil feedbacks on the growth of eastern North American trees: Implications for climate-adaptive forestry. Frontiers in Ecology and Evolution, 11. PDF

Royo, A. A., Raymond, P., Kern, C. C., Adams, B. T., Bronson, D., Champagne, E., Dumais, D., Gustafson, E., Marquardt, P. E., McGraw, A. M., Miesel, J. R., Munson, A. D., Périé, C., Tavares-Moreira, F. J., Ola, A., Bouchard, M., & Bissonnette, J. F. (2023). Desired REgeneration through Assisted Migration (DREAM): Implementing a research framework for climate-adaptive silviculture. Forest Ecology and Management, 546, 121298. PDF

Spicer, M. E., Royo, A. A., Wenzel, J. W., & Carson, W. P. (2023). Understory plant growth forms respond independently to combined natural and anthropogenic disturbances. Forest Ecology and Management, 543, 121077. PDF

Stout, S. L., Hille, A. T., & Royo, A. A. (2023). Science-management collaboration is essential to address current and future forestry challenges. In C. C. Kern & Y. L. Dickinson (Eds.), Proceedings of the first biennial Northern Hardwood Conference 2021: Bridging science and management for the future (Gen. Tech. Rep. NRS-P-211, pp. 15–24). U.S. Department of Agriculture, Forest Service, Northern Research Station. PDF

Yacucci, A. C., Carson, W. P., Martineau, J. C., et al. (2023). Native tree species prosper while exotics falter during gap-phase regeneration, but only where deer densities are near historical levels. New Forests. PDF

2022

Kramer, D. W., Prebyl, T. J., Nibbelink, N. P., Miller, K. V., Royo, A. A., & Frair, J. L. (2022). Managing moose from home: Determining landscape carrying capacity for Alces alces using remote sensing. Forests, 13(2), 150. PDF

Nieves, J. M., Steiner, K. C., McDill, M. E., Kreye, J. K., Ward, J. S., & Royo, A. A. (2022). Stand and site characteristics affect the probability of stump sprouting in some eastern North American hardwoods. Forest Ecology and Management, 509, 120136. PDF

Reed, S. P., Royo, A. A., Fotis, A. T., Knight, K. S., Flower, C. E., & Curtis, P. E. (2022). The long-term impacts of deer herbivory in determining temperate forest stand and canopy structure. Journal of Applied Ecology, 59(3), 812–821. PDF

Royo, A. A., & Carson, W. P. (2022). Stasis in forest regeneration following deer exclusion and understory gap creation: A ten-year experiment. Ecological Applications, 32(4), e2569. PDF

2021

Champagne, E., Raymond, P., Royo, A. A., Speed, J. D. M., Tremblay, J. P., & Côté, S. D. (2021). A review of ungulate impacts on the success of climate-adapted forest management strategies. Current Forestry Reports, 7, 305–320. PDF

Champagne, E., Royo, A. A., Tremblay, J. P., & Raymond, P. (2021). Tree assisted migration in a browsed landscape: Can we predict susceptibility to herbivores? Forest Ecology and Management, 498, 119576. PDF

Royo, A. A., Vickers, L. A., Long, R. P., Ristau, T. E., Stoleson, S. H., & Stout, S. L. (2021). The forest of unintended consequences: Anthropogenic actions trigger the rise and fall of black cherry. BioScience, 71(7), 683–696. PDF

2020

Champagne, E., Bonin, M., Royo, A. A., Tremblay, J. P., & Raymond, P. (2020). Predicting terpene content in dried conifer shoots using near infrared spectroscopy. Journal of Near Infrared Spectroscopy, 28, 308–314. PDF

Champagne, E., Royo, A. A., Tremblay, J. P., & Raymond, P. (2020). Phytochemicals involved in plant resistance to leporids and cervids: A systematic review. Journal of Chemical Ecology, 46(1), 84–98. PDF

De Vriendt, L., Thiffault, N., Royo, A. A., Barrette, M., & Tremblay, J. P. (2020). Moose browsing tends spruce plantations more efficiently than mechanical release. Forests, 11, 1138. PDF

Pinchot, C. C., Royo, A. A., Schlarbaum, S. E., Peters, M. P., Sharp, A. M., & Anagnostakis, S. L. (2020). The effect of site quality on performance of American chestnut (Castanea dentata) seedlings bred for blight (Cryphonectria parasitica) resistance. In C. D. Nelson, J. L. Koch, & R. A. Sniezko (Eds.), Proceedings of the Sixth International Workshop on the Genetics of Host-Parasite Interactions in Forestry (Gen. Tech. Rep. SRS–252). U.S. Department of Agriculture Forest Service, Southern Research Station.

Ristau, T. E., & Royo, A. A. (2020). Influence of stand age, soil moisture, and cover type on Rubus seed bank abundance. Journal of the Torrey Botanical Society, 147(3), 222–231. PDF

2019

Champagne, E., Royo, A. A., Tremblay, J. P., & Raymond, P. (2019). Tree Assisted Migration in a Browsed Landscape: Developing an Assessment of Susceptibility. In C. C. Kern & Y. L. Dickinson (Eds.), Proceedings of the First Biennial Northern Hardwood Conference 2021: Bridging science and management for the future (Gen. Tech. Rep. NRS-P-211, pp. 275–279). U.S. Department of Agriculture, Forest Service, Northern Research Station. https://doi.org/10.2737/NRS-GTR-P-211-paper58

McWilliams, W. H., Westfall, J. A., Brose, P. H., Dey, D. C., D’Amato, A. W., Dickinson, Y. L., Fajvan, M. A., Kenefic, L. S., Kern, C. C., Laustsen, K. M., Lehman, S. L., Morin, R. S., Ristau, T. E., Royo, A. A., Stoltman, A. M., & Stout, S. L. (2019). Subcontinental-scale patterns of large-ungulate herbivory and synoptic review of restoration management implications for midwestern and northeastern forests. Gen. Tech. Rep. NRS-182. U.S. Department of Agriculture Forest Service, Northern Research Station. https://doi.org/10.2737/NRS-GTR-182

Morgan, Q., Johnstone-Yellin, T., Pinchot, C. C., Peters, M., & Royo, A. A. (2019). Partitioning and predicting biomass to assess deer browse in Allegheny hardwood forests. Canadian Journal of Forest Research, 49(3), 301–316. PDF

Royo, A. A., Pinchot, C., Stanovick, J. S., & Stout, S. L. (2019). Timing is not everything: Assessing whether pre-versus post-harvest herbicide applications mitigate the burgeoning birch phenomenon in regenerating forests. Forests, 10(4), 324. PDF

Royo, A. A., & Stanovick, J. S. (2019). Deer browsing overwhelms extended leaf phenology benefits: A test case with Rubus allegheniensis and a recalcitrant hay-scented fern layer. Forest Ecology and Management, 448, 294–299. PDF

Slyder, J. B., Wenzel, J. W., Royo, A. A., Spicer, M. E., & Carson, W. P. (2019). Post-windthrow salvage logging increases seedling and understory diversity with little impact on composition immediately after logging. New Forests. PDF

Stout, S. L., Brose, P. H., Cleveland, H., Long, R. P., McGuinness, B., Peters, M. P., Rebbeck, J., Ristau, T. E., Royo, A. A., Stoleson, S. H., Thomasma, S., Twery, M. J., & Wurzbacher, S. (2019). Fifty years of science-management cooperation from the SILVAH community of practice. In S. L. Stout (Ed.), SILVAH: 50 years of science-management cooperation (Gen. Tech. Rep. NRS-P-186, pp. 8–25). U.S. Department of Agriculture Forest Service, Northern Research Station. PDF

Royo, A. A., & Stout, S. L. (2019). Recognition, response and recovery: Deer impact research in Allegheny hardwood forests. In S. L. Stout (Ed.), SILVAH: 50 years of science-management cooperation (Gen. Tech. Rep. NRS-P-186, pp. 26–36). U.S. Department of Agriculture Forest Service, Northern Research Station. PDF

Vickers, L. A., McWilliams, W. H., Knapp, B. O., D’Amato, A. W., Dey, D. C., Dickinson, Y. L., Kabrick, J. M., Kenefic, L. S., Kern, C. C., Larsen, D. R., Royo, A. A., Saunders, M. R., Shifley, S. R., & Westfall, J. A. (2019). Are current seedling demographics poised to regenerate northern US forests? Journal of Forestry, 93(1), 1–21. PDF

2018

Flower, C. E., Fant, J. B., Hoban, S., Knight, K. S., Steger, L., Aubihl, E., Gonzalez-Meler, M. A., Forry, S., Hille, A., & Royo, A. A. (2018). Optimizing conservation strategies for a threatened tree species: In situ conservation of White Ash (Fraxinus americana L.) genetic diversity through insecticide treatment. Forests, 9(4), 202. https://doi.org/10.3391/f9040202

Leverkus, A. B., Rey Benayas, J. M., Castro, J., Boucher, D., Brewer, S., Collins, B. M., Donato, D. C., Fraver, S., Kishchuk, B. E., Lee, E. J., Lindenmayer, D., Macdonald, E., Marzano, R., Rhoades, C. C., Royo, A. A., Thorn, S., Wagenbrenner, J. W., Waldron, K., Wohlgemuth, T., & Gustafsson, L. (2018). Salvage logging effects on regulating and supporting ecosystem services – A systematic map. Canadian Journal of Forest Research, 48(9), 983–1000. https://doi.org/10.1139/cjfr-2018-0114

McWilliams, W. H., Westfall, J. A., Brose, P. H., Dey, D. C., D’Amato, A. W., Dickinson, Y. L., Fajvan, M. A., Kenefic, L. S., Kern, C. C., Laustsen, K. M., Lehman, S. L., Morin, R. S., Ristau, T. E., Royo, A. A., Stoltman, A. M., & Stout, S. L. (2018). Subcontinental-scale patterns of large-ungulate herbivory and synoptic review of restoration management implications for midwestern and northeastern forests. Gen. Tech. Rep. NRS-182. USDA Forest Service, Northern Research Station. https://doi.org/10.2737/NRS-GTR-182

Raymond, P., Royo, A. A., Prévost, M., & Dumais, D. (2018). Assessing the single-tree and small group selection cutting system as intermediate disturbance to promote regeneration and diversity in temperate mixedwood stands. Forest Ecology and Management, 430, 21–32. https://doi.org/10.1016/j.foreco.2018.07.054

2017

Averill, K. M., Mortensen, D. A., Smithwick, E. A. H., Kalisz, S., McShea, W. J., Bourg, N. A., Parker, J. D., Royo, A. A., Abrams, M. D., Apsley, D. K., Blossey, B., Boucher, D. H., Caraher, K. L., DiTommaso, A., Johnson, S. E., Masson, R., & Nuzzo, V. A. (2017). A regional assessment of white-tailed deer effects on plant invasion. AoB Plants, 9, plx047. https://doi.org/10.1093/aobpla/plx047

Barrette, M., Bélanger, L., De Grandpré, L., & Royo, A. A. (2017). Demographic disequilibrium caused by canopy gap expansion and recruitment failure triggers forest cover loss. Forest Ecology and Management, 401, 117–124. https://doi.org/10.1016/j.foreco.2017.07.012

De Jager, N. R., Drohan, P. J., Miranda, B. M., Sturtevant, B. R., Stout, S. L., Royo, A. A., Gustafson, E. J., & Romanski, M. C. (2017). Simulating ungulate herbivory across forest landscapes: A browsing extension for LANDIS-II. Ecological Modelling, 350, 11–29. https://doi.org/10.1016/j.ecolmodel.2017.01.014

Flower, C. E., Aubihl, E., Fant, J., Forry, S., Hille, A., Knight, K. S., Oldland, W. K., Royo, A. A., & Turcotte, R. M. (2017). In-situ genetic conservation of white ash (Fraxinus americana) at the Allegheny National Forest. In R. A. Sniezko, G. Man, V. Hipkins, K. Woeste, D. Gwaze, J. T. Kliejunas, & B. A. McTeague (Tech. Cords.), Gene conservation of tree species—banking on the future (Gen. Tech. Rep. PNW-GTR-963, pp. 165–169). USDA Forest Service, Pacific Northwest Research Station. https://doi.org/10.2737/PNW-GTR-963

Kern, C. C., Burton, J. I., Raymond, P., D’Amato, A. W., Keeton, W. S., Royo, A. A., Walters, M. B., Webster, C. R., & Willis, J. L. (2017). Challenges in gap-based silviculture/forest management: Problems, potential solutions. Forestry, 90(1), 4–17. https://doi.org/10.1093/forestry/cpw024

McWilliams, W. H., Westfall, J. A., Brose, P. H., Lehman, S. L., Morin, R. S., Ristau, T. E., Royo, A. A., & Stout, S. L. (2017). After a quarter century, what does the Pennsylvania regeneration study tell us about the future of mixed oak forests under stress? In Proceedings of the 20th Central Hardwood Forest Conference (Gen. Tech. Rep. NRS-P-167, pp. 280–290). USDA Forest Service, Northern Research Station.

Royo, A. A., Kramer, D. W., Nibbelink, N. P., Miller, K. V., & Stout, S. L. (2017). Spatio-temporal variation in foodscapes modifies deer browsing impact on vegetation. Landscape Ecology, 32(12), 2281–2295. https://doi.org/10.1007/s10980-017-0568-x

Sinacore, K., Hall, J. S., Potvin, C., Royo, A. A., Ducey, M. J., & Ashton, M. S. (2017). Unearthing the hidden world of roots: Root biomass and architecture differ among species within the same guild. PLoS ONE, 12(10), e0185934. https://doi.org/10.1371/journal.pone.0185934

2016

Pendergast, T., Hanlon, S., Long, Z., Royo, A. A., & Carson, W. P. (2016). The legacy of deer overabundance: Long-term delays in wildflower community recovery. Canadian Journal of Forest Research, 46, 362–369. https://doi.org/10.1139/cjfr-2015-0280

Royo, A. A., Kramer, D. W., Nibbelink, N., Miller, K. V., & Stout, S. L. (2016). The canary in the coal mine: On the utility of sprouts as a rapid indicator of browse impact in disturbed forests. Ecological Indicators, 69, 269–275. https://doi.org/10.1016/j.ecolind.2016.04.030

Royo, A. A., Peterson, C. J., Stanovick, J., & Carson, W. P. (2016). Evaluating the ecological impacts of salvage logging: Can natural and anthropogenic disturbances promote coexistence? Ecology, 97(7), 1566–1582. https://doi.org/10.1890/15-1093.1

2015

Avella, A., Cárdenas, L. M., Torres, S., & Royo, A. A. (2015). Restoration of oak forests in the Colombian Andes. In J. A. Stanturf & P. Madsen (Eds.), Restoration of Boreal and Temperate Forests (2nd ed.). CRC Press.

Chips, M. J., Yerger, E. H., Havernak, A., Nuttle, T., Royo, A. A., Pruitt, J. N., McGlynn, T. P., Riggall, C. L., & Carson, W. P. (2015). The indirect impact of long-term overbrowsing on arthropods in the Allegheny National Forest region of Pennsylvania. Northeastern Naturalist, 22(4), 782–797. https://doi.org/10.1656/045.022.0412

McWilliams, W. H., Westfall, J. A., Brose, P. H., Dey, D. C., Hatfield, M., Johnson, K., Laustsen, K. M., Lehman, S., Morin, R. S., Nelson, M. D., Ristau, T. E., Royo, A. A., Stout, S. L., Willard, T., & Woodall, C. W. (2015). A regeneration indicator for Forest Inventory and Analysis: History, sampling, estimation, analytics, and potential use in the Midwest and Northeast U.S. (Gen. Tech. Rep. NRS-148). USDA Forest Service, Northern Research Station. https://doi.org/10.2737/NRS-GTR-148

Rooney, T. P., Madsen, P., Buttenschon, R., Olesen, C. R., Royo, A. A., & Stout, S. L. (2015). Integrating ungulate herbivory into forest landscape restoration. In J. A. Stanturf & P. Madsen (Eds.), Restoration of Boreal and Temperate Forests (2nd ed.). CRC Press.

2014

Carson, W. P., Royo, A. A., & Peterson, C. J. (2014). A pox on our land: A case study of chronic deer overbrowsing throughout the Allegheny National Forest Region of Pennsylvania. In F. S. Gilliam & M. R. Roberts (Eds.), The Herbaceous Layer in Forests of Eastern North America (2nd ed., pp. 400–411). Oxford University Press.

Huebner, C. D., Steinman, J., Hutchinson, T. F., Ristau, T. E., & Royo, A. A. (2014). The distribution of a non-native (Rosa multiflora) and native (Kalmia latifolia) shrub in mature closed-canopy forests across soil fertility gradients. Plant and Soil, 377(1–2), 259–276. https://doi.org/10.1007/s11104-013-2000-x

Knight, K. S., & Royo, A. A. (2014). Searching high and low: Patterns of white ash health across topographic gradients in the Allegheny region (Project NE-EM-09-02). In K. M. Potter & B. L. Conkling (Eds.), Forest Health Monitoring: National Status, Trends, and Analysis 2012 (Gen. Tech. Rep. SRS-198, pp. 161–166). USDA Forest Service.

Nuttle, T., Ristau, T., & Royo, A. A. (2014). Long-term biological legacies of herbivore density in a landscape-scale experiment: Forest understories reflect past deer density treatments for at least 20 years. Journal of Ecology, 102, 221–228. https://doi.org/10.1111/1365-2745.12175

2013

Nuttle, T., Royo, A. A., Adams, M. B., & Carson, W. P. (2013). Historic disturbance regimes promote tree diversity only under low browsing regimes in eastern deciduous forests. Ecological Monographs, 83(1), 3–17. https://doi.org/10.1890/11-2263.1

Peterson, C. J., Royo, A. A., Krueger, L., Stark, S. A., & Carson, W. P. (2013). Disturbance size and severity covary in small and mid-size wind disturbances in Pennsylvania northern hardwood forests. Forest Ecology and Management, 302, 273–279. https://doi.org/10.1016/j.foreco.2013.03.045

Royo, A. A., & Ristau, T. E. (2013). Stochastic and deterministic processes regulate spatio-temporal variation in seed bank diversity. Journal of Vegetation Science, 24, 724–734. https://doi.org/10.1111/jvs.12011

Stout, S. L., Royo, A. A., deCalesta, D. S., McAleese, K., & Finley, J. C. (2013). The Kinzua Quality Deer Cooperative: Can adaptive management and local stakeholder engagement sustain reduced impact of ungulate browsers in forest systems? Boreal Environment Research, 18(Suppl. A), 50–64.

2012

Dey, D. C., Royo, A. A., Gardiner, E. S., & Cardenas, L. M. (2012). Sustaining Quercus humboldtii and Colombobalanus excelsa on the Colombian landscape: Preservation or conservation—a research perspective. In J. R. Butnor (Ed.), Proceedings of the 16th Biennial Southern Silvicultural Research Conference (pp. 88–90). USDA Forest Service, e-GTR-SRS-156.

Huebner, C. D., Hutchinson, T., Ristau, T., Royo, A. A., & Steinman, J. (2012). Documenting the regional and local distribution of Kalmia latifolia and Rosa multiflora in West Virginia, Ohio, and Pennsylvania forests along a soil fertility gradient. In K. M. Potter & B. L. Conkling (Eds.), Forest health monitoring: 2009 national technical report (Gen. Tech. Rep. SRS-167, pp. 211–217). USDA Forest Service, Southern Research Station.

Royo, A. A., & Knight, K. S. (2012). White ash (Fraxinus americana) decline and mortality: The role of site nutrition and stress history. Forest Ecology and Management, 286, 8–15. https://doi.org/10.1016/j.foreco.2012.08.049

Royo, A. A., Knight, K. S., Himes, J. M., & Will, A. N. (2012). White ash (Fraxinus americana) health in the Allegheny Plateau Region, Pennsylvania: Evaluating the relationship between FIA Phase 3 crown variables and a categorical rating system. In W. McWilliams & F. A. Roesch (Eds.), Monitoring Across Borders: 2010 Joint Meeting of the Forest Inventory and Analysis (FIA) Symposium and the Southern Mensurationists (e-Gen. Tech. Rep. SRS-157, p. 299). USDA Forest Service, Southern Research Station.

2011

Goetsch, C., Wigg, J., Royo, A. A., Ristau, T., & Carson, W. P. (2011). Chronic overbrowsing and biodiversity collapse in the Allegheny Plateau Region, PA: Evidence from a 60-year-old deer exclosure. Journal of the Torrey Botanical Society, 138(2), 220–224.

Kain, M., Battaglia, L., Royo, A. A., & Carson, W. P. (2011). Over-browsing in Pennsylvania creates a depauperate forest dominated by an understory tree: Results from a 60-year-old deer exclosure. Journal of the Torrey Botanical Society, 138(3), 322–326.

Royo, A. A., Heartsill Scalley, T., Moya, S., & Scatena, F. N. (2011). Non-arborescent vegetation trajectories following repeated hurricane disturbance: Ephemeral versus enduring responses. Ecosphere, 2(7). https://doi.org/10.1890/ES11-0018.1

2010

Dey, D. C., Royo, A. A., Brose, P. H., Hutchinson, T. F., Spetich, M. A., & Stoleson, S. H. (2010). An ecologically based approach to oak silviculture: A synthesis of 50 years of oak ecosystem research in North America. Revista Colombia Forestal, 13, 5–30.

Knight, K. S., Herms, D. A., Cardina, J., Long, R., Rebbeck, J., Gandhi, J. J. K., Smith, A., Klooster, W. S., Herms, C. P., & Royo, A. A. (2010). Emerald ash borer aftermath forests: The dynamics of ash mortality and the responses of other plant species. In C. H. Michler & M. D. Ginzel (Eds.), Proceedings of the symposium on ash in North America, March 9–11, 2010 (Gen. Tech. Rep. NRS-P-72). USDA Forest Service, Northern Research Station.

Reinhart, K., Royo, A. A., Kageyama, S. A., & Clay, K. (2010). Canopy gaps decrease microbial densities and disease risk for a shade-intolerant tree species. Acta Oecologica, 36, 530–536. https://doi.org/10.1016/j.actao.2010.07.006

Royo, A. A., Carson, W. P., Collins, R., Adams, M. B., & Kirschbaum, C. (2010). Pervasive interactions between keystone ungulates and historical disturbance regimes promote temperate forest herbaceous diversity. Ecology, 91, 93–105. PDF

Royo, A. A., Carson, W. P., & Stout, S. L., Decalesta, D. S., & Pierson, T. (2010). Restoring forest herb communities through landscape-level deer herd reductions: Is recovery limited by legacy effects? Biological Conservation, 143, 2425–2434. https://doi.org/10.1016/j.biocon.2010.05.020

Royo, A. A., & Carson, W. P. (2010). On the formation of dense understory layers in forests worldwide: Consequences and implications for forest dynamics, biodiversity, and succession. In J. M. Pye, H. M. Rauscher, Y. Sands, D. C. Lee, & J. S. Beatty (Eds.), Advances in Threat Assessment and their Application to Forest and Rangeland Management (PNR-GTR-802, pp. 469–498). USDA Forest Service.

2009

Krueger, L. M., Peterson, C. J., Royo, A. A., & Carson, W. P. (2009). Evaluating relationships among tree growth rate, shade tolerance, and browse tolerance following disturbance in an eastern deciduous forest. Canadian Journal of Forest Research, 39, 2460–2469. https://doi.org/10.1139/X09-155

2008

Royo, A. A., & Carson, W. P. (2008). Direct and indirect effects of a dense understory on tree seedling recruitment in temperate forests: Habitat-mediated predation versus competition. Canadian Journal of Forest Research, 38, 1634–1645. https://doi.org/10.1139/X07-247

Royo, A. A., Bates, R., & Lacey, E. P. (2008). Demographic constraints in three populations of Lobelia boykinii: A rare wetland endemic. Journal of the Torrey Botanical Society, 135(2), 189–199.

2006

Royo, A. A., & Carson, W. P. (2006). On the formation of dense understory layers in forests worldwide: Consequences and implications for forest dynamics, biodiversity, and succession. Canadian Journal of Forest Research, 36, 1345–1362. PDF

2005

Banta, J. A., Royo, A. A., Kirschbaum, C., & Carson, W. P. (2005). Evidence for the ghost of herbivory past: Plant community refugia on boulders in the Allegheny National Forest. Natural Areas Journal, 25(1), 10–18.

Comisky, L., Royo, A. A., & Carson, W. P. (2005). Deer browsing creates rock refugia gardens on large boulders in the Allegheny National Forest, Pennsylvania. American Midland Naturalist, 154, 201–206. https://doi.org/10.1674/0003-0031(2005)154[0201:DBCRRG]2.0.CO;2

Reinhart, K. O., Royo, A. A., Van der Putten, W. H., & Clay, K. (2005). Soil feedback and pathogen activity in Prunus serotina throughout its native range. Journal of Ecology, 93, 890–898. https://doi.org/10.1111/j.1365-2745.2005.01028.x

Royo, A. A., & Carson, W. P. (2005). The herb community of a tropical forest in central Panamá: Dynamics and impact of mammalian herbivores. Oecologia, 145(1), 66–75. https://doi.org/10.1007/s00442-005-0079-3

2001

Lacey, E. P., Royo, A. A., Bates, R., & Herr, D. (2001). The role of population dynamic models in biogeographic studies: Illustrations from a study of Lobelia boykinii. Castanea, 66(1–2), 115–125.